Ithete, Ndapewa Laudika; Stoffberg, Samantha; Corman, Victor Max; และคณะ (2013). "Close Relative of Human Middle East Respiratory Syndrome Coronavirus in Bat, South Africa". Emerging Infectious Diseases. 19 (10): 1697–1699. doi:10.3201/eid1910.130946. ISSN1080-6040.
Lau, Susanna K P; Zhang, Libiao; Luk, Hayes K H; และคณะ (2018). "Receptor Usage of a Novel Bat Lineage C Betacoronavirus Reveals Evolution of Middle East Respiratory Syndrome-Related Coronavirus Spike Proteins for Human Dipeptidyl Peptidase 4 Binding". The Journal of Infectious Diseases. 218 (2): 197–207. doi:10.1093/infdis/jiy018. ISSN0022-1899.
Ithete, Ndapewa Laudika; Stoffberg, Samantha; Corman, Victor Max; และคณะ (2013). "Close Relative of Human Middle East Respiratory Syndrome Coronavirus in Bat, South Africa". Emerging Infectious Diseases. 19 (10): 1697–1699. doi:10.3201/eid1910.130946. ISSN1080-6040.
Lau, Susanna K P; Zhang, Libiao; Luk, Hayes K H; และคณะ (2018). "Receptor Usage of a Novel Bat Lineage C Betacoronavirus Reveals Evolution of Middle East Respiratory Syndrome-Related Coronavirus Spike Proteins for Human Dipeptidyl Peptidase 4 Binding". The Journal of Infectious Diseases. 218 (2): 197–207. doi:10.1093/infdis/jiy018. ISSN0022-1899.