Blanco, D.R.; Lui, R.L.; Vicari, M.R.; Bertollo, L.A.C.; Moreira-Filho, O. (2011). "Comparative Cytogenetics of Giant Trahiras Hoplias aimara and H. intermedius (Characiformes, Erythrinidae): Chromosomal Characteristics of Minor and Major Ribosomal DNA and Cross-Species Repetitive Centromeric Sequences Mapping Differ among Morphologically Identical Karyotypes". Cytogenetic and Genome Research. 132 (1–2): 71–78. doi:10.1159/000320923. PMID20924165.
Blanco, D.R.; Lui, R.L.; Vicari, M.R.; Bertollo, L.A.C.; Moreira-Filho, O. (2011). "Comparative Cytogenetics of Giant Trahiras Hoplias aimara and H. intermedius (Characiformes, Erythrinidae): Chromosomal Characteristics of Minor and Major Ribosomal DNA and Cross-Species Repetitive Centromeric Sequences Mapping Differ among Morphologically Identical Karyotypes". Cytogenetic and Genome Research. 132 (1–2): 71–78. doi:10.1159/000320923. PMID20924165.