Brenner, Don J.; Krieg, Noel R.; Staley, James T. (26 Temmuz 2005) [1984(Williams & Wilkins)]. George M. Garrity (Ed.). The Proteobacteria. Bergey's Manual of Systematic Bacteriology. 2C (2. bas.). New York: Springer. ss. 1388. ISBN978-0-387-24145-6. British Library no. GBA561951.