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Hekkala, E.; Shirley, M.H.; Amato, G.; Austin, J.D.; Charter, S.; Thorbjarnarson, J.; Vliet, K.A.; Houck, M.L.; Desalle, R.; Blum, M.J. (2011). An ancient icon reveals new mysteries: Mummy DNA resurrects a cryptic species within the Nile crocodile. Molecular Ecology. 20 (20): 4199—4215. doi:10.1111/j.1365-294X.2011.05245.x. PMID21906195.
Robert W. Meredith, Evon R. Hekkala, George Amato and John Gatesy (2011). A phylogenetic hypothesis for Crocodylus (Crocodylia) based on mitochondrial DNA: Evidence for a trans-Atlantic voyage from Africa to the New World. Molecular Phylogenetics and Evolution. 60 (1): 183—191. doi:10.1016/j.ympev.2011.03.026. PMID21459152.