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Verdú, José R.; Galante, Eduardo; Lumaret, Jean-Pierre; Cabrero-Sañudo, Francisco J. (2004). Phylogenetic analysis of Geotrupidae (Coleoptera, Scarabaeoidea) based on larvae. Systematic Entomology. 29 (4): 509—523. doi:10.1111/j.0307-6970.2004.00256.x. ISSN0307-6970.(англ.)