Herron, Matthew D.; Castoe, Todd A.; Parkinson, Christopher L. (2004). Sciurid phylogeny and the paraphyly of Holarctic ground squirrels (Spermophilus). Molecular Phylogenetics and Evolution. 31 (3): 1015—30. doi:10.1016/j.ympev.2003.09.015. PMID15120398.
Helgen, K. M.; Cole, F. R.; Helgen, L. E.; Wilson, D. E. Generic revision in the Holarctic ground squirrel genus Spermophilus // Journal of Mammalogy. — 2009. — Вип. 90. — № 2. — С. 270–305. — DOI:10.1644/07-MAMM-A-309.1.(англ.)
Herron, Matthew D.; Castoe, Todd A.; Parkinson, Christopher L. (2004). Sciurid phylogeny and the paraphyly of Holarctic ground squirrels (Spermophilus). Molecular Phylogenetics and Evolution. 31 (3): 1015—30. doi:10.1016/j.ympev.2003.09.015. PMID15120398.