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Saccheri, I.; Kuussaari, M.; Kankare, M.; Vikman, P.; Fortelius, W.; Hanski, I. (1998). Inbreeding and extinction in a butterfly metapopulation. Nature. 392 (6675): 491. Bibcode:1998Natur.392..491S. doi:10.1038/33136.
Laine, Anna-Liisa (2016). Ilkka Hanski (1953–2016) Population ecologist who modelled how species cope with habitat loss. Nature. 534 (7606): 180. doi:10.1038/534180a. PMID27279205.