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Aschliman; Nishida; Miya; Inoue; Rosana; Naylord (2012). Body plan convergence in the evolution of skates and rays (Chondrichthyes: Batoidea). Molecular Phylogenetics and Evolution. 63 (1): 28—42. doi:10.1016/j.ympev.2011.12.012. PMID22209858.
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