Patterson, S.A.; Morris-Pocock, J.A.; Friesen, V.L (2011). "A multilocus phylogeny of the Sulidae (Aves: Pelecaniformes)". Molecular Phylogenetics and Evolution. Quyển 58 số 2. tr. 181–191. doi:10.1016/j.ympev.2010.11.021. PMID21144905.
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pubmed.ncbi.nlm.nih.gov
Patterson, S.A.; Morris-Pocock, J.A.; Friesen, V.L (2011). "A multilocus phylogeny of the Sulidae (Aves: Pelecaniformes)". Molecular Phylogenetics and Evolution. Quyển 58 số 2. tr. 181–191. doi:10.1016/j.ympev.2010.11.021. PMID21144905.