David P. Maitland (1992). “Carapace movements aid air breathing in the semaphore crab, Heloecius cordiformis (Decapoda: Brachyura: Ocypodidae)”. Journal of Comparative Physiology B. 162: 375–382. doi:10.1007/BF00260766.
D. R. Fielder; J. G. Greenwood (1985). “The systematic position of Heloecius cordiformis (H. Milne Edwards, 1837) (Decapoda, Ocypodidae) as revealed by larval morphology”. Crustaceana. 48 (3): 244–248. doi:10.1163/156854085X00954. JSTOR20104041.
jstor.org
D. R. Fielder; J. G. Greenwood (1985). “The systematic position of Heloecius cordiformis (H. Milne Edwards, 1837) (Decapoda, Ocypodidae) as revealed by larval morphology”. Crustaceana. 48 (3): 244–248. doi:10.1163/156854085X00954. JSTOR20104041.