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Haldane, J. B. S. (1990). “A mathematical theory of natural and artificial selection--I. 1924”. Bulletin of Mathematical Biology. 52 (1–2): 209–40, discussion 201–207. doi:10.1016/s0092-8240(05)80010-2. PMID2185859. Originally published in Trans Camb Phil Soc 23: 19–41.