M. E. Laidre & R. W. Elwood (2008). “Motivation matters: cheliped extension displays in the hermit crab, Pagurus bernhardus, are honest signals of hunger”. Animal Behaviour. 75 (6): 2041–2047. doi:10.1016/j.anbehav.2007.11.011.
Brian A. Hazlett (1967). “Interspecific shell fighting between Pagurus bernhardus and Pagurus cuanensis (Decapoda, Paguridea)”. Sarsia. 29 (1): 215–220. doi:10.1080/00364827.1967.10411083.