Hemmer, H.; Kahlike, R.D.; Vekua, A.K. (2004). “The Old World puma Puma pardoides (Owen, 1846) (Carnivora: Felidae) in the Lower Villafranchian (Upper Pliocene) of Kvabebi (East Georgia, Transcaucasia) and its evolutionary and biogeographical significance”. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 233: 197–233. doi:10.1127/njgpa/233/2004/197.