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Stelbrink, B.; von Rintelen, T.; Cliff, G.; Kriwet, J. Molecular systematics and global phylogeography of angel sharks (genus Squatina). Molecular Phylogenetics and Evolution. 2010, 54 (2): 395–404. PMID 19647086. doi:10.1016/j.ympev.2009.07.029.
Nagasawa, K.; Tanaka, S.; Benz, G.W. Trebius shiinoi n. sp. (Trebiidae: Siphonostomatoida: Copepoda) from uteri and embryos of the Japanese angelshark (Squatina japonica) and the clouded angelshark (Squatina nebulosa), and redescription of Trebius longicaudatus. Journal of Parasitology. 1998, 84 (6): 1218–1230. PMID 9920318. doi:10.2307/3284678.