Ward, Philip S.; Brady, Sean G.; Fisher, Brian L.; Schultz, Ted R. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology. July 2014, 40 (1): 61–81. ISSN 1365-3113. doi:10.1111/syen.12090.
Ward, Philip S.; Brady, Sean G.; Fisher, Brian L.; Schultz, Ted R. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology. July 2014, 40 (1): 61–81. ISSN 1365-3113. doi:10.1111/syen.12090.