Ishiguro, Naotaka; Inoshima, Yasuo; Shigehara, Nobuo; Ichikawa, Hideo; Kato, Masaru. Osteological and Genetic Analysis of the Extinct Ezo Wolf (Canis Lupus Hattai) from Hokkaido Island, Japan. Zoological Science. 2010, 27 (4): 320–324. PMID 20377350. doi:10.2108/zsj.27.320.
Matsumura, Shuichi; Inoshima, Yasuo; Ishiguro, Naotaka. Reconstructing the colonization history of lost wolf lineages by the analysis of the mitochondrial genome. Molecular Phylogenetics and Evolution. 2014, 80: 105–112. PMID 25132126. doi:10.1016/j.ympev.2014.08.004.
Koblmüller, Stephan; Vilà, Carles; Lorente-Galdos, Belen; Dabad, Marc; Ramirez, Oscar; Marques-Bonet, Tomas; Wayne, Robert K.; Leonard, Jennifer A. Whole mitochondrial genomes illuminate ancient intercontinental dispersals of grey wolves (Canis lupus). Journal of Biogeography. 2016, 43 (9): 1728–1738. doi:10.1111/jbi.12765.
Ishiguro, Naotaka; Inoshima, Yasuo; Shigehara, Nobuo; Ichikawa, Hideo; Kato, Masaru. Osteological and Genetic Analysis of the Extinct Ezo Wolf (Canis Lupus Hattai) from Hokkaido Island, Japan. Zoological Science. 2010, 27 (4): 320–324. PMID 20377350. doi:10.2108/zsj.27.320.
Matsumura, Shuichi; Inoshima, Yasuo; Ishiguro, Naotaka. Reconstructing the colonization history of lost wolf lineages by the analysis of the mitochondrial genome. Molecular Phylogenetics and Evolution. 2014, 80: 105–112. PMID 25132126. doi:10.1016/j.ympev.2014.08.004.