Churcher, C.S.; De Iuliis, G. A new species of Protopterus and a revision of Ceratodus humei (Dipnoi: Ceratodontiformes) from the Late Cretaceous Mut Formation of eastern Dakhleh Oasis, Western Desert of Egypt. Palaeontology(英语:Palaeontology (journal)). 2001, 44 (2): 305–323. S2CID 128764735. doi:10.1111/1475-4983.00181.
Churcher, C.S.; De Iuliis, G. A new species of Protopterus and a revision of Ceratodus humei (Dipnoi: Ceratodontiformes) from the Late Cretaceous Mut Formation of eastern Dakhleh Oasis, Western Desert of Egypt. Palaeontology(英语:Palaeontology (journal)). 2001, 44 (2): 305–323. S2CID 128764735. doi:10.1111/1475-4983.00181.
Churcher, C.S.; De Iuliis, G. A new species of Protopterus and a revision of Ceratodus humei (Dipnoi: Ceratodontiformes) from the Late Cretaceous Mut Formation of eastern Dakhleh Oasis, Western Desert of Egypt. Palaeontology(英语:Palaeontology (journal)). 2001, 44 (2): 305–323. S2CID 128764735. doi:10.1111/1475-4983.00181.