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Witmer, L.M., S. Chatterjee, J. Franzosa, T. Rowe, and R. C. Ridgely. (2004). "Neuroanatomy and vestibular apparatus of pterosaurs: Implications for flight, posture, and behavior." Annual Meeting of the Society of Integrative and Comparative Biology, New Orleans, LA. Integrative and Comparative Biology, 43(6): 832. [1] (页面存档备份,存于互联网档案馆)
Witmer, L.M., S. Chatterjee, J. Franzosa, T. Rowe, and R. C. Ridgely. (2004). "Neuroanatomy and vestibular apparatus of pterosaurs: Implications for flight, posture, and behavior." Annual Meeting of the Society of Integrative and Comparative Biology, New Orleans, LA. Integrative and Comparative Biology, 43(6): 832. [1] (页面存档备份,存于互联网档案馆)