Campbell, J.; J. Vannini. A new subspecies of beaded lizard, Heloderma horridum, from the Motagua Valley of Guatemala. Journal of Herpetology. 1988, 22 (4): 457–468. JSTOR 1564340. doi:10.2307/1564340.
Campbell, J.; J. Vannini. A new subspecies of beaded lizard, Heloderma horridum, from the Motagua Valley of Guatemala. Journal of Herpetology. 1988, 22 (4): 457–468. JSTOR 1564340. doi:10.2307/1564340.