Wiklund, Christer; Tullberg, Birgitta S. Seasonal polyphenism and leaf mimicry in the comma butterfly. Animal Behaviour. September 2004, 68 (3): 621–627. S2CID 54270418. doi:10.1016/j.anbehav.2003.12.008.
Endler, John A. An Overview of the Relationships Between Mimicry and Crypsis. Biological Journal of the Linnean Society. August 1981, 16 (1): 25–31. doi:10.1111/j.1095-8312.1981.tb01840.x.
Wiklund, Christer; Tullberg, Birgitta S. Seasonal polyphenism and leaf mimicry in the comma butterfly. Animal Behaviour. September 2004, 68 (3): 621–627. S2CID 54270418. doi:10.1016/j.anbehav.2003.12.008.