Schubart, Christoph D.; Cuesta, José A.; Felder, Darryl L. GLYPTOGRAPSIDAE, A NEW BRACHYURAN FAMILY FROM CENTRAL AMERICA: LARVAL AND ADULT MORPHOLOGY, AND A MOLECULAR PHYLOGENY OF THE GRAPSOIDEA. Journal of Crustacean Biology. 2002-02, 22 (1): 28–44. doi:10.1651/0278-0372(2002)022[0028:GANBFF]2.0.CO;2.
Schubart, C. D.; Cannicci, S.; Vannini, M.; Fratini, S. Molecular phylogeny of grapsoid crabs (Decapoda, Brachyura) and allies based on two mitochondrial genes and a proposal for refraining from current superfamily classification. Journal of Zoological Systematics and Evolutionary Research. 2006-08, 44 (3): 193–199. doi:10.1111/j.1439-0469.2006.00354.x.
Schubart, Christoph D.; Liu, Hung-Chang & Cuesta, José A. (2003): A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bulletin of Zoology51(1): 49-59. PDF fulltext (页面存档备份,存于互联网档案馆)
Schubart, Christoph D.; Liu, Hung-Chang & Cuesta, José A. (2003): A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bulletin of Zoology51(1): 49-59. PDF fulltext (页面存档备份,存于互联网档案馆)