Bogdanov, AS; Bannikova, AA; Pirusskiĭ, IuM; Formozov, NA. The first genetic evidence of hybridization between west European and northern white-breasted hedgehogs (Erinaceus europaeus and E. Roumanicus) in Moscow Region. Izvestiia Akademii nauk. Seriia biologicheskaia / Rossiiskaia akademiia nauk. 2009, (6): 760–5. PMID 20143638.