Hiroaki Aiba and David Král. 2023. The First Fossil of Ceratophyus (Coleoptera, Geotrupidae) from the Middle Pleistocene Shiobara Group of Nasushiobara City, Tochigi Prefecture, Japan. Paleontological Research. 28(3); 265-272. DOI: https://doi.org/10.2517/PR230013
