Rhodin, A.G.J.; Thomson, S.; Georgalis, G.; Karl, H.-V.; Danilov, I.G.; Takahashi, A.; de la Fuente, M.S.; Bourque, J.R.; Delfino M.; Bour, R.; Iverson, J.B.; Shaffer, H.B.; van Dijk, P.P. Turtles and tortoises of the world during the rise and global spread of humanity: first checklist and review of extinct Pleistocene and Holocene chelonians. Chelonian Research Monographs 5. 2015: 000e.1–66. ISBN 978-0965354097. doi:10.3854/crm.5.000e.fossil.checklist.v1.2015.
Adán Pérez-García; Evangelos Vlachos; Xabier Murelaga. A large testudinid with African affinities in the post-Messinian (lower Pliocene) record of south-eastern Spain. Palaeontology. 2020, 63 (3): 497–512. S2CID 214232312. doi:10.1111/pala.12468.
Adán Pérez-García; Evangelos Vlachos; Xabier Murelaga. A large testudinid with African affinities in the post-Messinian (lower Pliocene) record of south-eastern Spain. Palaeontology. 2020, 63 (3): 497–512. S2CID 214232312. doi:10.1111/pala.12468.