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Kellner, Alexander W. A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renan A. M.; Sayão, Juliana M. A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert. Anais da Academia Brasileira de Ciências. August 19, 2019, 91 (suppl 2): e20190768. ISSN 0001-3765. PMID 31432888. doi:10.1590/0001-3765201920190768(英语).
Holgado, B.; Pêgas, R.V. A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae. Acta Palaeontologica Polonica. 2020, 65. doi:10.4202/app.00751.2020.
Kellner, Alexander W. A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renan A. M.; Sayão, Juliana M. A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert. Anais da Academia Brasileira de Ciências. August 19, 2019, 91 (suppl 2): e20190768. ISSN 0001-3765. PMID 31432888. doi:10.1590/0001-3765201920190768(英语).
Kellner, Alexander W. A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renan A. M.; Sayão, Juliana M. A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert. Anais da Academia Brasileira de Ciências. August 19, 2019, 91 (suppl 2): e20190768. ISSN 0001-3765. PMID 31432888. doi:10.1590/0001-3765201920190768(英语).