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Ozaki, Kenichi; Isono, Masahiro; Kawahara, Takayuki; Iida, Shigeo; Kudo, Takuma; Fukuyama, Kenji (2006). "A Mechanistic Approach to Evaluation of Umbrella Species as Conservation Surrogates". Conservation Biology. 20 (5): 1507–1515. doi:10.1111/j.1523-1739.2006.00444.x. PMID17002768. S2CID8383041.